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Kasugano Haruka and his sister Sora have lost both their parents in an accident, and with them all their support. They decide to move out of the city to the rural town where they once spent summers with their late grandfather. At first everything seems familiar and peaceful, but changes come as Haruka starts to remember things from his youth.

After the foods win their war against the humans at the end of the film, they create Cibopolis, a city of their own ruled entirely by foods. But soon, problems arise, including rain, crows, and internal discord among the foods. To find answers, Frank, Brenda, and Barry set out to find a human.